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Costa Rica, December 2018

Posted: Fri Dec 21, 2018 4:05 pm
by petesmith
I have just returned from a wonderful two week holiday in the Guanacaste province of Costa Rica, in the far north west of the country, not far from the Nicaraguan border. We stayed in a remote hotel on the coast, surrounded by tropical forests full of wildlife such as Howler Monkeys, Spider Monkeys, Anteaters, Iguanas, and all manner of other creatures. Guanacaste is the driest region of Costa Rica, and we were there at the beginning of the dry season, but the vegetation was still very lush and green.

I should stress at this point that this was not a butterflying holiday, but given our surroundings it was inevitable that some interesting species would turn up, and as this was my first time out of Europe, I was like a kid in a sweetshop! Of course, there was the added bonus that every butterfly species seen would be a life tick...

We spent much of the fortnight in and around our hotel, taking a daily early morning walk of a couple of hours along the tracks through the jungle before the heat forced us back to the pool for cocktails and some R&R. I was glad to have with me a copy of Jeffrey Glassberg's excellent book, "A Swift Guide to Butterflies of Mexico and Central America". This book manages to pack the maximum amount of information on a large number of species into a relatively small field guide and is very cleverly designed and laid out. Even so, there were some species that I struggled to identify, and I shall be requesting assistance from anyone with experience of central American butterflies in due course! Also, if anyone disagrees with any of my tentative IDs I would value any imput.

The following species were all photographed within a couple of kilometres of our base, at elevations between sea-level and 50 metres:

I didn't have to wait long on our first morning for the excitement of seeing my first new non-European butterflies. A walk through the hotel gardens produced the following three delights.
White Peacock, Anartia jatrophae2.JPG
White Peacock - Anartia jatrophae
Banded Peacock, Anartia fatima.JPG
Banded Peacock - Anartia fatima
Pale-banded Crescent, Phyciodes tulcis2.JPG
Pale-banded Crescent, Phyciodes tulcis unds.JPG
Pale-banded Crescent - Phyciodes tulcis

The tracks through the forest were alive with butterflies - I didn't know where to point my eyes first! One of the species to catch my attention early was the diminutive Elf butterfly, which was flying in good numbers along the side of the track. They are delightful insects, constantly opening and closing their wings, and quite active.
Elf, Microtia elva2.JPG
Elf, Microtia elva unds.JPG
Elf - Microtia elva

There were also some tiny blues moving about. I eventually managed a few photographs and identified them as Ceraunus Blues - they wouldn't look completely out of place in parts of Europe.
Ceraunus Blue, Hemiargus ceraunus5.JPG
Ceraunus Blue, Hemiargus ceraunus4.JPG
Ceraunus Blue, Hemiargus ceraunus3.JPG
Next up was one of the metalmarks, a group of butterflies that here in Europe we seem to have been short-changed on. I think this is a Bright Scintillant, another tiny butterfly no larger than a female common blue, but with exquisitely beautiful markings. The taxonomy of these beasts is still uncertain, and Glassberg doesn't even offer a species name for this one.
Bright Scintillant, Calephelis sp..JPG
Bright Scintillant - Calephelis sp

A little further along, the forest track opened up into open grassland, and along the edge were a number of small fritillary-like butterflies, soon to become a personal favourite of mine, with stunning underside patterns and rather nice uppers also! These were Theona Checkerspots.
Theona Checkerspot, Chlosyne theona3.JPG
Theona Checkerspot, Chlosyne theona pair2.JPG
Theona Checkerspot, Chlosyne theona.JPG
Theona Checkerspot - Chlosyne theona

But hang on a minute, we were in the tropics here, and so far all the butterflies we had seen had been pretty small. Shouldn't there be some larger ones flying about? This thought had just gone through my mind when a huge Morpho flew through the trees alongside us. It wasn't stopping for a picture but it was a stunning sight as it flew low down and flashed its irridescent blue at us several times on its way past!
Over the coming days, this forest track became our regular morning haunt, and produced more and more species of interest.
The Malachite was commonly seen in ones and twos, usually up in the tree tops, but occasionally descending lower down. It took me a whole week to get a photo of one at nectar without severe wing damage, as most of them were way past their best.
Malachite, Siproeta stelenes.JPG
Malachite - Siproeta stelenes
Another sizeable butterfly, although not quite as big as a Monarch, was the Soldier.
Soldier, Danaus eresimus10.JPG
Soldier - Danaus eresimus


There was a good selection of whites and sulphurs on the wing, but getting anywhere near them was not easy, as they were mostly restless. I should mention that there was a stiff breeze blowing most days, so even if a butterfly did land briefly closeby, it was often challenging to get a camera on it. Below are a few shots of varying quality.
Tailed Orange, Eurema proterpia.JPG
Tailed Orange - Eurema proterpia
Boisduval's Yellow, Eurema boisduvaliana.JPG
Boisduval's Yellow - Eurema boisduvaliana
Yellow-and-white Yellow, Eurema venusta.JPG
Yellow-and-white Yellow - Eurema venusta
Cloudless Sulphur, Phoebis sennae.JPG
Cloudless Sulphur - Phoebis sennae

One morning we walked upon a huge emergence of Barred Yellows - they were everywhere, flying over and around some vetch plants, with many mating pairs - the previous day we hadn't seen a single one.
Barred Yellow, Eurema daira.JPG
Barred Yellow, Eurema daira 2.JPG
Barred Yellow - Eurema daira


I had hoped to bump into one of the Buckeyes during our visit, and I wasn't disappointed. These were fairly common.
Tropical Buckeye, Junonia genoveva.JPG
Tropical Buckeye - Junonia goneveva

The next species was a complete revelation. I had read about "Cracker" butterflies, but seeing them in the wild doing their thing was amazing. As two rival males interact and fight over territories, they make a surprisingly loud "cracking" noise as part of their battle, which can be clearly heard from some metres away.The sound has been likened to that of a firecracker going off, but to me it was reminiscent of the sound of a high voltage electric discharge, like the sound that accompanies a Van der Graaf Generator spark. At first I thought that these were the Variable Cracker, Hamadryas feronia, but on closer examination I believe this is the Guatemalan Cracker.
Guatemalan Cracker, Hamadryas guatemalena3.JPG
Guatemalan Cracker, Hamadryas guatemalena2.JPG
Guatemalan Cracker - Hamadryas guatemalena

A larger narrow-winged fritillary had been eluding me for a few days before I was able to get close enough to photograph the gorgeous Mexican Fritillary.
Mexican Fritillary, Euptoieta hegesia.JPG
Mexican Fritillary, Euptoieta hegesia unds.JPG
Mexican Fritillary - Euptoieta hegesia

This one also caused me a few headaches to begin with - quite appropriately - which Sister was this? I am fairly confident of this ID now.
Confusing Sister, Adelpha iphicleola2.JPG
Confusing Sister - Adelpha iphicleola

Swallowtails were surprisingly thin on the ground, and the only species we saw was this one.
Giant Swallowtail, Papilio cresphontes.JPG
Giant Swallowtail - Papilio cresphontes

Another species that appeared out of nowhere one morning was the Black Patch. We saw these on just one day, in small numbers.
Black Patch, Chlosyne melanarge pair.JPG
Black Patch - Chlosyne melanarge

And to finish this first part of my report, a species which according to Glassberg is endemic to Mexico, so I have either got the ID wrong, or this butterfly also flies in NW Costa Rica. I have this down as a worn example of the White-tipped Pixie, but stand ready to be corrected!
White-tipped Pixie, Melanis acroleuca.JPG
White-tipped Pixie - Melanis acroleuca

In PART TWO I shall share my experience with the Skipper butterflies that we encountered. I think they really were the highlight of the trip for me, and I am not known as a huge skipper fan usually! There was a fascinating range, from the "near familiar" ones that closely resemble some of our European Skippers, through to the "otherworldly" the likes of which you could scarcely imagine!

Re: Costa Rica, December 2018

Posted: Fri Dec 21, 2018 8:26 pm
by petesmith
PART TWO
The Skippers

I shall start with those that I am fairly confident I have identified, and move onto some which I would welcome some help with!
The first Skippers that I found seemed almost familiar, not too far removed from some of the European Pyrgus species. By far the commonest in our local area was the Tropical Checkered Skipper.
Tropical Checkered Skipper, Pyrgus oileus3.JPG
Tropical Chequered Skipper - Pyrgus oileus
Very similar, but much less common, was the close relative, Common Chequered Skipper.
Common Chequered Skipper, Pyrgus communis.JPG
Common Chequered Skipper - Pyrgus communis
Another species with a European feel to it appeared on the wing about half way through our holiday:
Whirlabout, Polites vibex.JPG
Whirlabout, Polites vibex unds.JPG
Whirlabout - Polites vibex

Below is the Brazilian Skipper.
Brazilian Skipper, Calpodes ethlius.JPG
Brazilian Skipper - Calpodes ethlius
Manuel's Skipper was only seen on one occasion, but posed nicely at nectar before zipping off at high speed.
Manuel's Skipper, Polygonus manueli2.JPG
Manuel's Skipper, Polygonus manueli.JPG
Manuel's Skipper - Polygonus manueli

After this, things began to get a bit more exotic, as the following selection shows. There was a damp mud-puddling hole just a few metres from the beach, on the edge of the forest, and it seemed that a different species was in residence every morning, either mud-puddling, or just sat around nearby.
White Enops, Polyctor cleta.JPG
White Enops - Polyctor cleta
White-patched Skipper, Chiomara asychis2.JPG
White-patched Skipper, Chiomara asychis unds.JPG
White-patched Skipper - Chiomara asychis
Common Bentwing, Ebriatas anacreon.JPG
Common Bentwing - Ebrietas anacreon
Glassy-winged Skipper, Xenophanes tryxus@.JPG
Glassy-winged Skipper - Xenophanes tryxus
Dark-spotted Polythrix, Polythrix asine.JPG
Dark-spotted Polythrix, Polythrix asine4.JPG
Dark-spotted Polythrix - Polythrix asine
Double-striped Longtail, Urbanus belli.JPG
Double-striped Longtail - Urbanus belli
Pale Mylon, Mylon pelopidas2.JPG
Pale Mylon - Mylon pelopidas

The Dusted Spurwing below is one of my favourite photographs from our trip - I love the wing textures on this beauty!
Dusted Spurwing, Antigonus erosus.JPG
Dusted Spurwing - Antigonus erosus

And the next species looks very unlike any skipper I have ever seen before. This one was common along the forest track edges and in more open areas. Another absolutely gorgeous insect.
Veined White-Skipper, Heliopetes arsalte2.JPG
Veined White-Skipper, Heliopetes arsalte.JPG
Veined White-skipper - Heliopetes arsalte


qChocolate-marked Therra, Thespius dalman.JPG
qChocolate-marked Therra, Thespius dalman.JPG
Laviana White-skipper - Helioptes laviana/i]
qDun Skipper, Euphyes vestris.JPG
Dun Skipper - Euphyes vestris
qFritzgaertner's Flat, Celaenorrhinus fritzgaertneri.JPG
Fritzgaertner's Flat - Celaenorrhinus fritzgaertneri - this one flew into our hotel room late one afternoon!
qMountain Scallopwing, Staphylus vincula@.JPG
Mountain Scallopwing - Staphylus vincula - this was taken at higher altitude around 600m asl further east in Costa Rica, close to the continental divide, in much damper cloud forest habitat.
qSkipper 1.JPG
Any ideas on this anyone?

Part three to follow, with a couple of hairstreak queries and a few other photos from inland Costa Rica.

Re: Costa Rica, December 2018

Posted: Fri Dec 21, 2018 9:40 pm
by bugboy
Hi Pete, this webpage might be useful. Hover over each species name and you get an image straight away, beats clicking on name after name when you have a species list like Costa Rica! :)

http://www.tropicleps.ch/?page=4

Re: Costa Rica, December 2018

Posted: Fri Dec 21, 2018 10:06 pm
by petesmith
Thanks bugboy! I shall check this out tomorrow, when hopefully my jetlag will have finally disappeared!

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 12:57 pm
by David M
Thanks for brightening up a winter day for me, Pete! Spellbinding array of species there already and I dread to know how many images you took...I suspect well into four figures!

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 1:40 pm
by Tony Moore
Hi Pete,

C.R. really is a spellbinding place, especially for butterflyers, and should be on everyone's 'bucket' list.

I,too, thought the 'Crackers' quite amazing insects although the 'cracking' was too high pitched for my old lugs. When I tried to photograph them in poor light (where they're usually found), the pre-flash caused them to flip their wings and spoil the pic. I spent some time trying to chase them onto tree trunks in more light. All part of the fun :mrgreen: .

Merry Christmas all,

Tony M.

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 3:23 pm
by petesmith
David - you are correct! I came back with 1274 images to process, which is actually quite restrained, but less than half of these were butterflies. There was just so much else to point the cameras at!

Tony - yes, the Crackers always settled on tree trunks, always facing downwards, and usually in shady places some way up the trunks, so it wasn't that easy to get photo's! But perseverance paid off in the end...

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 3:56 pm
by petesmith
PART THREE
Here are a final few photographs from our trip.
Hairstreaks were few and far between, and only two came within reach of the camera lens, but they were a joy to see:
Clytie Ministreak, Ministrymon clytie.JPG
Clytie Ministreak - Ministrymon clytie
Emessa Hairstreak, Gargina emessa.JPG
Emessa Hairstreak - Gargina emessa

Below is the Tabasco Yellow.
Tabasco Yellow, Eurema agave.JPG
Tabasco Yellow - Eurema agave

And another Pierid that initially proved challenging to identify:
Lyside Sulphur, Kricogonia lyside.JPG
Lyside Sulphur - Kricogonia lyside

We spent three days away from our hotel, exploring many of the attractions further east in Costa Rica. Here we were close to the continental divide and actually crossed over to the Caribbean half of the country on a couple of occasions. The habitat was much damper, with cloud forest and volcanoes, and we were at much higher altitude. A few different species were to be found, but as we were often in cloud, little was flying.
Rusty-tipped Page, Siptoeta epaphus@.JPG
Rusty-tipped Page - Siptoeta epaphus
Dot-bordered Heliconian, Heliconius doris@.JPG
Dot-bordered Heliconian - Heliconius doris
Carolina Satyr, Hermeuptychia sosybius.JPG
Carolina Satyr - Hermeuptychia sosybus
Common Morpho, Morpho helenor.JPG
Common Morpho, Morpho helenor unds.JPG
Common Morpho - Morpho helenor

And finally, a species that I have been completely unable to identify. I am starting to wonder if it is actually a butterfly...
qWhat is this@.JPG
This truly was the trip of a lifetime, and the butterflies were just a small part of the incredible range of wildlife we encountered. Sloths, Coatis, Anteaters, Bats, Frogs, Lizards, Birds and other insects, stunning scenery, wonderful people. I can strongly recommend Costa Rica as a destination to any wildlife enthusiast!

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 6:14 pm
by bugboy
Your last one looks like another member of the moth family Castniinae. I say another as there was a recent thread that caused much interweb searching with a related species

viewtopic.php?f=14&t=10035&p=139518#p139518

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 7:44 pm
by petesmith
Thanks again bugboy - I am now happy with the moth Castniomera drucei as an ID for this one. To be honest it was behaving rather strangely for a butterfly!
I have also now changed the ID of my ?Chocolate -marked Therra to Laviana White-skipper - Helioptes laviana

Your help is much appreciated!

Re: Costa Rica, December 2018

Posted: Sat Dec 22, 2018 9:27 pm
by Charles Nicol
thanks for sharing those lovely images Pete !

Re: Costa Rica, December 2018

Posted: Sun Dec 23, 2018 12:46 am
by Sooty
Some great butterflies there! I'm spending 18 days in Huatulco, Mexico in February and I plan on spending a few days getting out with my camera.

Re: Costa Rica, December 2018

Posted: Sun Dec 23, 2018 6:51 pm
by petesmith
Charles - it is a pleasure to share images of such beauties!

Sooty - best of luck in Mexico - I look forward to seeing what you are able to find out there!

Re: Costa Rica, December 2018

Posted: Sun Dec 23, 2018 8:06 pm
by David M
Looks like you've visited a butterfly house, Pete.....but of course, you haven't; you saw these in the wild.

Sumptuous.

Re: Costa Rica, December 2018

Posted: Sun Dec 23, 2018 8:22 pm
by petesmith
I know what you mean David - it was a case of seeing things that you have only ever seen in Zoos before! Just strolling down the local track and an Anteater casually crosses your path...monkeys hanging out of the local trees...sloths also. And the butterflies would not look out of place in a tropical butterfly house - it was fantastic seeing them, and the other local flora and fauna in their natural environment. A very special experience...

Re: Costa Rica, December 2018

Posted: Thu Dec 27, 2018 12:47 pm
by Medard
Hello Pete, a great collection of photos from your trip, a reward for the boring nature of a long haul flight. Brilliant ! you mentioned seeing a coati,or coatimundi it triggered memories of a trip my wife and I made to Geronimo territory Chiricahua National Monument, Arizona,where we were lucky to see one. The park rangers identified it and told us, "you Guys are very lucky they are very elusive critters".
Thanks for the memory.

Re: Costa Rica, December 2018

Posted: Thu Dec 27, 2018 4:36 pm
by Tony Moore
When we were in Costa Rica a white-nosed Coati regularly dug up the lawn in front of one of our hotels. It was totally unfazed by humans and allowed close approach. A magic experience :) .
DSC08318.JPG
.

Tony M.

Re: Costa Rica, December 2018

Posted: Thu Dec 27, 2018 9:41 pm
by petesmith
Jim, Tony - the Coati was indeed a wonderful experience, as were many of the close encounters we had with a whole range of exotic wildlife, and thanks Jim for your kind comments on my photos.

This guy below was pretty cool, just strolling down the road towards me without a care in the world:
Coati 2.JPG
I think my mind is still in Costa Rica...it takes some getting over!
But I have now tried to enter the "post-analytical" phase after the holiday, going through photos and records and re-living the whole experience.

To keep this thread on track, after a couple of vertebrate pics (of which I could add lots more, but that would be very self-indulgent and off topic!), here is another shot of one butterfly that I have been researching since arriving home.

It is the White-tipped Pixie, Melanis acroleuca, said by Glassberg to be a Mexican endemic, but one that I photographed in NW Costa Rica.
White-tipped Pixie, Melanis acroleuca2.JPG
I have since found out that this butterfly has other names, such as white-rayed metalmark, and also possibly other specific names such as Melanis cephise, which may or may not be synonymous with acroleuca.

Taxonomy is never simple is it! Especially when you are more than 5000 miles away from home...

The distribution maps certainly seem to have it as an almost exclusively Mexican species, although there is at least one Costa Rican record on inaturalist, albeit with a less than convincing photo...

Re: Costa Rica, December 2018

Posted: Sat Dec 29, 2018 10:43 pm
by Charles Nicol
petesmith wrote:here is another shot of one butterfly that I have been researching since arriving home.

It is the White-tipped Pixie, Melanis acroleuca, said by Glassberg to be a Mexican endemic, but one that I photographed in NW Costa Rica.

White-tipped Pixie, Melanis acroleuca2.JPG

I have since found out that this butterfly has other names, such as white-rayed metalmark, and also possibly other specific names such as Melanis cephise, which may or may not be synonymous with acroleuca.

Taxonomy is never simple is it! Especially when you are more than 5000 miles away from home...

The distribution maps certainly seem to have it as an almost exclusively Mexican species, although there is at least one Costa Rican record on inaturalist, albeit with a less than convincing photo...
it looks like the White-tipped and White-rayed are different species of Melanis:

the 4th & 5th pics here should help

http://www.martinreid.com/Butterfly%20w ... marks.html

Re: Costa Rica, December 2018

Posted: Sun Dec 30, 2018 12:04 am
by bugboy
Here's a couple of species accounts that may aid, both with reasonably detailed distribution maps.

https://www.inaturalist.org/taxa/474547 ... -acroleuca

https://www.inaturalist.org/taxa/148508-Melanis-cephise